By Robert L. Dressler, Kerry Dressler
As but, a accomplished ordinary category of the orchids isn't really to be had, due partly to their bewildering variety and parallelism, and likewise to botanical forget. even supposing a lot paintings is required ahead of a completely traditional category should be completed, this publication offers a useful contribution to the on-going discussions. The author's transparent demeanour of writing, and his said services during this topic, make the bankruptcy on category of particular curiosity. His paintings delimits numerous significant traditional teams and emphasises parts wanting additional learn, relatively one of the extra primitive teams. info on orchid chromosome numbers is introduced jointly and a brand new interpretation of this information supplied. Novel info on pollination can be awarded. a listing of wide-spread names and demanding synonyms is given, in addition to approximate numbers of species in every one genus. Dr Dressler has been provided the 1994 Henry Allan Gleason Award via the hot York Botanical backyard for Phylogeny and category of the Orchid kin. The award is gifted each year for a good booklet within the fields of taxonomy, plant ecology, and plant geography.
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Extra info for Phylogeny and Classification of the Orchid Family
Rasmussen (1986a) notes, however, that the stipe of Stanhopea is several cell layers thick. 38 39 In some other orchids, a similar structure develops in a different way. In the Tropidieae, Prasophyllinae, and Bulbophyllum ecornutum, for example, an extension of the rostellum forms a connection between the pollinia and the viscidium. This is termed a "hamulus," or hamular stipe. POLARITY. Each type of stipe has evolved independently in several groups, and caudicles are usually a prerequisite for the evolution of stipes (not in the Bulbophyllinae).
Rather than evolving a single median viscidium, this group evolved two distinct viscidia, one attached to each of the pollinia, with the rostellar extension between them. Some Orchideae have a single viscidium, but even these usually have the rostellar extension above the viscidium, thus suggesting that the single viscidium is secondary in this group. In Orchis and Dactylorhiza each viscidium is covered by a sheath, or bursicle, that is easily broken when touched, thus exposing the fresh, sticky viscidium.
Sensitive Rostellum In Listera, the rostellum includes a sack, or pocket, of glue under pressure. When a pollinator touches the rostellum this sack breaks open, squirting out fresh glue onto the intruder. At the same time, the margins of the rostellum reflex, releasing the pollinia to fall onto the fresh glue droplet (Ackerman and Mesler 1979; Schick et al. 1987). The sensitive rostellum of Listera may have evolved from a membrane-covered rostellum like that of Epipactis (Dressler 1990a). Accessory Structures More than the pollen mass, itself, is involved in the transfer of pollinia from one flower to another.